Stock Collection: 15,000 - 90,000 photos

General Subject Areas: Over 450 species of amphibians and reptiles, plus insects, spiders, birds, mammals, wildflowers, Mississippi River scenics, paddlewheelers, towboats and barges,

Books / Publication Credits: National Geographic, National Wildlife, Audubon, Natural History, Life, Sierra, Wildlife Conservation, Backpacker, Voyageur, Ranger Rick, Field & Stream, Sports Afield. Author/photographer of book: Amphibians & Reptiles of the North Woods.

Geographic Coverage: Wisconsin, Minnesota, Upper Mississippi River, Trempealeau NWR, Driftless Area, Everglades NP, Florida, Shawnee NF, Chequamegon NF, Perrot State Park, Great River Bluffs State Park,

Other services: Digital submissions available.

Typical Stock Photo Examples:
NATURAL HISTORY, protective coloration, camouflage, mimicry, warning coloration, salamanders migrating, frogs and toads calling, deformed frogs, deformed leopard frog, turtle laying eggs, turtle eggs hatching, turtle basking, lizard tail loss, skink tending eggs, lizard eggs hatching, snake eggs hatching, snake shedding skin, snake eating, venomous snakes, rattlesnakes at den, waterfowl on Mississippi River, bald eagles along Mississippi River, bald eagle flying, pelicans on Mississippi River, tundra swans on Mississippi River, egrets at Trempealeau NWR, Great Blue Heron at Trempealeau NWR, killdeer broken wing act, NATURAL HABITATS, bluffs, bluff prairies, oak savanna, vernal ponds, ponds, wetlands, rivers, marshes, swamps, forests, WILDLIFE MANAGEMENT, rare species, endangered species, radiotelemetry, bird banding, bird watching, frog crossing sign, turtle crossing sign, tortoise crossing sign, INSECTS, Mayfly, Mayflies, Dragonfly, Damselfly, Blue Dasher, Common Whitetail, Eastern Amberwing, Eastern Pondhawk, Twelve-spotted Skipper, Widow Skimmer, Walkingstick, Milkweed Bug, Linnaeus 17-year Cicada, Dobsonfly, Fishfly, Eyed Click Beetle, Long-horned Beetle, Red Milkweed Beetle, Tiger Beetle, Bee Fly, Hover Fly, Flower Fly, Mydas Fly, Robber Fly, Butterflies, Coral Hairstreak, Karner Blue, Karner Blue caterpillar, Monarch, Regal Fritillary, Tawny Emperor, Tiger Swallowtail, Viceroy, Broad-winged Skipper, Common Roadside Skipper, Delaware Skipper, Fiery Skipper, Little Glassywing, Mulberry Wing, Northern Broken Dash, Cecropia Moth, Hummingbird Moth, Isabella Moth, Lichen Moth, Net-winged Moth, Woolly Bear Moth, Polyphemus Moth, Primrose Moth, Sphinx Moth, Underwing Moth, Honey Bee, Bumble Bee, Cicada Killer, Paper Wasp, Yellow Jacket, Ants. ARACHNIDS, SPIDERS, Argiope, Black Widow, Crab Spiders, Fishing Spider, Northern Widow, Nursery Web Spider, Purseweb Spider, Wolf Spider, AMPHIBIANS, SALMANDERS, Aquatic Caecilian, Two-toed Amphiuma, One-toed Amphiuma, Eastern Hellbender, Alabama Waterdog, Mudpuppy, Dwarf Siren, Eastern Lesser Siren, Greater Siren, Ringed Salamander, Flatwoods Salamander, Jefferson Salamander, Blue-spotted Salamander, Spotted Salamander, Marbled Salamander, Mole Salamander, Smallmouth Salamander, Barred Tiger Salamander, Gray Tiger Salamander, Eastern Tiger Salamander, Broken-striped Newt, Red-spotted Newt, California Newt, Seepage Salamander, Apalachicola Dusky Salamander, Southern Dusky Salamander, Spotted Dusky Salamander, Northern Dusky Salamander, Imitator Salamander, Seal Salamander, Mountain Dusky Salamander, Blackbelly Salamander, Black Mountain Salamander, Pigmy Salamander, Southern Two-lined Salamander, Junaluska Salamander, Three-lined Salamander, Longtail Salamander, Dark-sided Salamander, Cave Salamander, Dwarf Salamander, Blue Ridge Two-lined Salamander, Blue Ridge Spring Salamander, Four-toed Salamander, Tellico Salamander, Redback Salamander, Red-backed Salamander, Slimy Salamander, Atlantic Coast Slimy Salamander, Eastern Zigzag Salamander, White-spotted Slimy Salamander, Northern Slimy Salamander, Southeastern Slimy Salamander, Mississippi Slimy Salamander, Jordans Salamander, Cumberland Plateau Salamander, Southern Redback Salamander, Southern Red-backed Salamander, Yonahlossee Salamander, Midland Mud Salamander, Rusty Mud Salamander, Northern Red Salamander, Blackchin Red Salamander, Southern Red Salamander, TOADS, FROGS, Firebelly Toad, Clawed Frog, African Clawed Frog, Spadefoot Toad, Couchs Spadefoot, Eastern Spadefoot Toad, Plains Spadefoot Toad, Horned Frog, Cranwells Horned Frog, Ornate Horned Frog, Greenhouse Frog, Budgett's Frog, Eastern American Toad, Dwarf American Toad, Boreal Toad, Great Plains Toad, Wyoming Toad, Eastern Green Toad, Western Green Toad, Giant Toad, Marine Toad, Southern Toad, Gulf Coast Toad, Texas Toad, European Green Toad, Fowlers Toad, Woodhouses Toad, Lesser Swimming Frog, Glass Frog, Dart Frog, Poison Frog, Green and Black Poison Frog, Blue Poison Frog, Yellow-banded Poison Frog, Strawberry Poison Frog, Dyeing Poison Frog, Blanchards Cricket Frog, Northern Cricket Frog, Florida Cricket Frog, Red-eyed Treefrog, Pine Barrens Treefrog, Bird-voiced Treefrog, Copes Gray Treefrog, Green Treefrog, Emerald Eye Treefrog, Pine Woods Treefrog, Barking Treefrog, Squirrel Treefrog, Gray Treefrog, Cuban Treefrog, Whites Treefrog, Dusky Treefrog, Mountain Chorus Frog, Brimleys Chorus Frog, Northern Spring Peeper, Ornate Chorus Frog, Upland Chorus Frog, Western Chorus Frog, Green and Golden Bell Frog, Bicolor Monkey Frog, Waxing Monkey Frog, Painted Monkey Frog, Greenhouse Frog, Solomon Island Leaf Frog, Painted Mantella, African Bullfrog, Florida Gopher Frog, Carolina Gopher Frog, Bullfrog, Bronze Frog, Green Frog, Red-eared Frog, Pig Frog, Florida Bog Frog, Pickerel Frog, Northern Leopard Frog, Mink Frog, Wood Frog, Southern Leopard Frog, Carpenter Frog, African Reed Frog, Spotted Reed Frog, Marbled Reed Frog, Spotted Running Frog, Amani Forest Treefrog, Clown Treefrog, Masked Big-eyed Frog, Big-eyed Forest Treefrog, Vietnamese Mossy Treefrog, Eastern Narrowmouth Toad, Great Plains Narrowmouth Toad, Malaysian Narrowmouth Toad, Muellers Narrowmouth Toad, Tomato Frog, Red-banded Crevice Creeper, South African Snake-neck Frog, REPTILES, TURTLES, Helmeted Turtle, Spot-bellied Side-neck Turtle, Giant Amazon River Turtle, Six-tubercled River Turtle, Yellow-spotted Amazon River Turtle, Matamata, Redbelly Shortneck Turtle, Common Snapping Turtle, Florida Snapping Turtle, Alligator Snapping Turtle, Striped Mud Turtle, Florida Mud Turtle, Yellow Mud Turtle, Big Bend Mud Turtle, Mississippi Mud Turtle, Eastern Mud Turtle, Razorback Musk Turtle, Loggerhead Musk Turtle, Stripeneck Musk Turtle, Stripe-necked Musk Turtle, Common Musk Turtle, Stinkpot, Western Painted Turtle, Southern Painted Turtle, Midland Painted Turtle, Eastern Painted Turtle, Reeves Turtle, Spotted Turtle, Wood Turtle, Western Pond Turtle, Bog Turtle, Southeast Asia Box Turtle, Keeled Box Turtle, Florida Chicken Turtle, Western Chicken Turtle, Eastern Chicken Turtle, Blandings Turtle, Barbours Map Turtle, Cagles Map Turtle, Yellow-blotched Map Turtle, Common Map Turtle, Northern Map Turtle, Black-knobbed Map Turtle, Ringed Map Turtle, Ouachita Map Turtle, Mississippi Map Turtle, False Map Turtle, Alabama Map Turtle, Giant Asian Pond Turtle, Carolina Diamondback Terrapin, Ornate Diamondback Terrapin, Mississippi Diamondback Terrapin, Northern Diamondback Terrapin, Japanese Turtle, Yellow Pond Turtle, Chinese Stripe-necked Turtle, Eastern River Cooter, Hieroglyphic River Cooter, Suwannee River Cooter, Florida Cooter, Peninsula Cooter, Redbelly Turtle, Red-bellied Turtle, Florida Red-bellied Turtle, American Redbelly Turtle, Indian Thorn Turtle, Painted Wood Turtle, Beals Eyed Turtle, Black Marsh Turtle, Florida Box Turtle, Eastern Box Turtle, Gulf Coast Box Turtle, Three-toed Box Turtle, Ornate Box Turtle, Sliders, Red-eared Slider, Yellowbelly Slider, Yellow-bellied Slider, Cumberland Slider, Softshell Turtles, Florida Softshell, Gulf Coast Smooth Softshell, Midland Smooth Softshell, Gulf Coast Spiny Softshell, Texas Spiny Softshell, Guadalupe Spiny Sofshell, Western Spiny Softshell, Eastern Spiny Softshell, Tortoises, South African Bowsprit Tortoise, Redfoot Tortoise, Indian Star Tortoise, Aldabra Tortoise, Leopard Tortoise, Radiated Tortoise, African Spurred Tortoise, Desert Tortoise, Gopher Tortoise, Elongated Tortoise, Bells Hingeback Tortoise, Homes Hingeback Tortoise, Asian Brown Tortoise, Burmese Brown Tortoise, African Pancake Tortoise, Central Asia Tortoise, LIZARDS, Florida Worm Lizard, Green Anole, Brown Anole, Mexican Spinytail Iguana, Collared Lizard, Desert Iguana, Texas Horned Lizard, Chuckwalla, Yellow-backed Spiny Lizard, Northern Plateau Lizard, Northern Prairie Lizard, Southern Fence Lizard, Rosebelly Lizard, Northern Side-blotch Lizard, Prairie Racerunner, Six-lined Racerunner, Western Whiptail, Northern Whiptail, Southern Coal Skink, Five-lined Skink, Southeastern Five-lined Skink, Broad-headed Skink, Broadhead Skink, Great Plains Skink, Northern Prairie Skink, Alligator Lizard, Legless Lizards, Western Slender Glass Lizard, Eastern Slender Glass Lizard, Island Glass Lizard, Eastern Glass Lizard, Gila Monster, Beaded Lizard, Veiled Chameleon, Usumbara Chameleon, Three-horn Chameleon, Bearded Dragon, Ornate Mastigure, Rough-scaled Plated Lizard, Banded Gecko, Leopard Gecko, Tokay Gecko, Yellow-headed Gecko, Brown Basilisk, Fiji Banded Iguana, Solomon Island Skink, Ocellated Skink, Bluetongue Skink, Sand Skink, Sandfish, Asian Grass Lizard, Savannah Monitor, Desert Monitor, Nile Monitor, Emerald Monitor, Roughneck Monitor, Chinese Crocodile Lizard, CROCODILIANS, American Alligator, Chinese Alligator, Spectacled Caiman, Yacare Caiman, Broad-snouted Caiman, Slender-snouted Crocodile, American Crocodile, Australian Freshwater Crocodile, Phillipine Crocodile, Morelets Crocodile, Mugger Crocodile, Cuban Crocodile, Gharial, Gavial, Black Caiman, Dwarf Crocodile, Dwarf Caiman, Smooth-fronted Caiman. SNAKES, Boa Constrictor, Burrowing Python, Rubber Boa, Emerald Tree Boa, Amazon Tree Boa, Cooks Tree Boa, Sand Boa, Rosy Boa, Yellow Anaconda, Green Anaconda, Green Tree Python, Blood Python, Burmese Python, Ball Python, Reticulate Python, Blind Snake, Glossy Snake, Worm Snake, Scarlet Snake, Northern Black Racer, Eastern Yellowbelly Racer, Eastern Yellow-bellied Racer, Prairie Ringneck Snake, Northern Ringneck Snake, Southern Ringneck Snake, Eastern Indigo Snake, Corn Snake, Great Plains Rat Snake, Texas Rat Snake, Black Rat Snake, Yellow Rat Snake, Everglades Rat Snake, Gray Rat Snake, Fox Snake, Eastern Mud Snake, Western Mud Snake, Western Hognose Snake, Eastern Hognose Snake, Southern Hognose Snake, Texas Night Snake, Gray-banded Kingsnake, Prairie Kingsnake, Mole Kingsnake, California Kingsnake, Eastern Kingsnake, Speckled Kingsnake, Sonoran Black Kingsnake, Desert Kingsnake, Durango Mountain Kingsnake, San Luis Potosi Kingsnake, Variable Kingsnake, Nuevo Leon Kingsnake, Louisiana Milk Snake, Mexican Milk Snake, Pueblan Milk Snake, New Mexico Milk Snake, Scarlet Kingsnake, Central Plains Milk Snake, Nelson's Milk Snake, Sinaloan Milk Snake, Red Milk Snake, Eastern Milk Snake, California Mountain Kingsnake, Arizona Mountain Kingsnake, Eastern Coachwhip, Western Coachwhip, Black Swamp Snake, Mangrove Salt Marsh Snake, Mississippi Green Water Snake, Redbelly Water Snake, Red-bellied Water Snake, Yellowbelly Water Snake, Yellow-bellied Water Snake, Copperbelly Water Snake, Copper-bellied Water Snake, Blotched Water Snake, Broad-banded Water Snake, Banded Water Snake, Diamondback Water Snake, Midland Water Snake, Northern Water Snake, Brown Water Snake, Rough Green Snake, Smooth Green Snake, Green Vine Snake, New Mexico Pine Snake, Sonora Gopher Snake, Northern Pine Snake, Bullsnake, Baja Cape Gopher Snake, Florida Pine Snake, Striped Crayfish Snake, Glossy Crayfish Snake, Queen Snake, Pine Woods Snake, Texas Longnose Snake, Black Swamp Snake, Ground Snake, Diadem Snake, Short-tailed Snake, Texas Brown Snake, Florida Redbelly Snake, Northern Redbelly Snake, Red-bellied Snake, Central Florida Crowned Snake, Butler's Garter Snake, Eastern Blackneck Garter Snake, Eastern Black-necked Garter Snake, Wandering Garter Snake, Checkered Garter Snake, Western Ribbon Snake, Eastern Plains Garter Snake, Western Plains Garter Snake, Bluestripe Ribbon Snake, Blue-striped Ribbon Snake, Peninsula Ribbon Snake, Red-sided Garter Snake, Bluestripe Garter Snake, Blue-striped Garter Snake, Common Garter Snake, Eastern Garter Snake, San Francisco Garter Snake, Lined Snake, Rough Earth Snake, Eastern Earth Snake, Sunbeam Snake, VENOMOUS SNAKES, POISONOUS SNAKES, Green Mamba, Eastern Coral Snake, Monocled Cobra, Black Pakistani Cobra, Red Spitting Cobra, Indian Cobra, Cape Cobra, Cantil, Florida Cottonmouth, Western Cottonmouth, Water Moccasin, Broad-banded Copperhead, Northern Copperhead, Southern Copperhead, Osage Copperhead, Trans-pecos Copperhead, Rough-scaled Bush Viper, Green Bush Viper, Puff Adder, Gaboon Viper, Rhinoceros Viper, Eyelash Palm Pit Viper, Urutu, Horned Viper, Eastern Diamondback Rattlesnake, Western Diamondback Rattlesnake, Sidewinder, Neotropical Rattlesnake, Aruba Island Rattlesnake, Timber Rattlesnake, Canebrake Rattlesnake, Banded Rock Rattlesnake, Mottled Rock Rattlesnake, Southwestern Speckled Rattlesnake, Panamint Rattlesnake, Blacktail Rattlesnake, Red Diamond Rattlesnake, Mojave Rattlesnake, Tiger Rattlesnake, Uracoan Rattlesnake, Arizona Black Rattlesnake, Midget Faded Rattlesnake, Northern Pacific Rattlesnake, Prairie Rattlesnake, Levantine Viper, Caucusus Viper, Saw-scaled Viper, Slender Hog-noed Pit Viper, Bushmaster, Eastern Massasauga, Western Massasauga, Dusky Pigmy Rattlesnake, Carolina Pigmy Rattlesnake, Western Pigmy Rattlesnake, Chinese Green Tree Viper, BIRDS, Pied-billed Grebe, Anhinga, Double-crested Cormorant, American White Pelican, Brown Pelican, Common Merganser, Hooded Merganser, Black Duck, Gadwall, Mallard, Northern Shoveler, Wood Duck, Blue-winged Teal, Ring-necked Duck, Canvasback, Lesser Scaup, Bluebills, Geese, Canada Goose, Trumpeter Swan, Tundra Swan, American Bittern, Herons, Egrets, Great Blue Heron, Great Egret, Green Heron, Little Blue Heron, Snowy Egret, Tricolored Heron, Yellow-crowned Night Heron, Black-crowned Night Heron, Ibis, White Ibis, Cranes, Sandhill Crane, Whooping Crane, Limpkin, Wood Stork, American Coot, Killdeer, Solitary Sandpiper, Greater Yellowlegs, Wilsons Phalarope, Long-billed Dowitcher, Common Nighthawk, Bald Eagle, American Kestrel, Peregrine Falcon, Cooper's Hawk, Red-shouldered Hawk, Red-tailed Hawk, Owls, Barred Owl, Boreal Owl, Burrowing Owl, Great Gray Owl, Great Horned Owl, Long-eared Owl, Northern Saw-whet Owl, Snowy Owl, Black Vulture, Turkey Vulture, Greater Prairie Chicken, Greater Prairie-Chicken, Sharp-tailed Grouse, American Woodcock, Downy Woodpecker, Red-bellied Woodpecker, Red-headed Woodpecker, American Tree Sparrow, Baltimore Oriole, Black-capped Chickadee, Blue Jay, Boat-tailed Grackle, Catbird, Dark-eyed Junco, Northern Cardinal, Red-winged Blackbird, Rusty Blackbird, Scrub Jay, Scarlet Tanager, Swainsons Thrush, Prothonotary Warbler, Yellow-rumped Warbler, Tree Swallow, Warbling Vireo, White-breasted Nuthatch, Yellow-headed Blackbird, MAMMALS, Opossum, RODENTS, Black-tailed Prairie Dog, Least Chipmunk, Gray Squirrel, Eastern Fox Squirrel, Eastern Gray Squirrel, Mouse, Deer Mouse, White-footed Mouse, Beaver, Muskrat, Porcupine, Eastern Cottontail, Marsh Rabbit, Snowshoe Hare, American Bison, Elk, Moose, White-tailed Deer, Mink, Marten, Fox, Arctic Fox, Common Gray Fox, Coyote, Gray Wolf, Timber Wolf, Arctic Fox, Red Fox, Silver Fox, Bobcat, Lynx, Mountain Lion, ALABAMA, Little River Canyon, FLORIDA, Wakulla Springs, Chinsegut Nature Center, Corkscrew Swamp, Sanibel, Anhinga Trail, ILLINOIS, Shawnee National Forest, Cache River State Natural Area, Heron Pond, La Rue-Pine Hills, IOWA, Pikes Peak State Park, Pike's Peak State Park, Effigy Mounds National Monument, Upper Mississippi River National Wildlife and Fish Refuge, Lansing, Guttenburg, Marquette, McGregor, MICHIGAN, Munising Falls, Miners Falls, Seney National Wildlife Refuge, MINNESOTA, Upper Mississippi River National Wildlife and Fish Refuge, Great River Bluffs State Park, Frontenac State Park, Beaver Creek Valley State Park, Whitewater State Park, Itasca State Park, Jay Cooke State Park, Banning State Park, Tettegouche State Park, Crow Wing State Park, Whitewater Wildlife Management Area, Kellogg-Weaver Dunes, Lake Winona, Barn Bluff, Colville Park, Reads Landing, Sugarloaf, Kings Bluff, Queens Bluff, Weaver Bottoms, Whitewater River, Uncas Dunes, Anoka Sand Plain, Winona, Red Wing, WISCONSIN, Upper Mississippi River National Wildlife and Fish Refuge, Amnicon State Park, Perrot State Park, Wyalusing State Park, Wildcat Mountain State Park, Nelson Dewey State Park, Devils Lake State Park, Devil's Lake State Park, Mill Bluff State Park, Natural Bridge State Park, Interstate Park, Roche-a-cri State Park, Trempealeau National Wildlife Refuge, Necedah National Wildlife Refuge, Crex Meadows Wildlife Area, Buena Vista Grasslands, Van Loon Wildlife Area, Riecks Lake, Sandhill Wildlife Management Area, Spring Green Preserve, Baxters Hollow, Parfreys Glen, The Hogback, Decorah Mound, St. Peters Dome, Cave Point County Park, Buena Vista Park, Memorial Park, Riverside Park, Bradys Bluff, Grandad Bluff, Battle Bluff, Rush Creek Bluff, Sugar Creek Bluff, Maiden Rock Bluff, Black River, Chippewa River, Red Cedar River, Namekagon River, Wolf River, Wisconsin River, Mississippi River, Tank Creek, St. Croix River, Kickapoo River, Amnicon River, Morgan Falls, Amnicon Falls, Hixon Forest, Goose Island County Park, Seven Bridges Trail, Great River Trail, Elroy-Sparta Trail, Great River Road, La Crosse, Arcadia, Alma, Cassville, Prairie du Chien, Nature Conservancy sites, UTAH, Arches National Monument, Brumley Creek, Mojave Desert, Snow Canyon, Zion National Park.,,,